What does 1 Chronicles 4:31 mean?

and at Beth-marcaboth, and Hazar-susim, and at Beth-biri, and at Shaaraim. These were their cities unto the reign of David.

And at Beth–marcaboth, and Hazar–susim, and at Beth–birei, and at Shaaraim. These were their cities unto the reign of David.

and at Beth-marcaboth, and Hazar-susim, and at Beth-biri, and at Shaaraim. These were their cities unto the reign of David.

And at Beth-marcaboth, and at Hazarsusim, and at Beth-biri, and at Shaaraim. These were their towns till David became king.

and in Beth-Marcaboth, and in Hazar-Susim, and in Beth-Birei, and in Shaarim; these <FI>are<Fi> their cities till the reigning of David.

And in Bethmarchaboth, and in Hasarsusim, and in Bethberai, and in Saarim. These were their cities unto the reign of David.

and at Beth-marcaboth, and Hazar-susim, and at Beth-biri, and at Shaaraim. These were their cities until the reign of David.